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Table 1 Lichen diversity and abundance on the architectural elements (arch. el.) in the site of the Tomb of Lazarus: antique churches (i), Monastery (ii), and modern church (iii). Growth forms of lichen species (crustose endolithic, Cr. en.; crustose epilithic, Cr. ep.; foliose, Fol.; squamulose, sq) and eutrophication index (1, not resistant to eutrophication; 2, resistant to a very weak eutrophication; 3, resistant to a weak eutrophication; 4, occurring in rather eutrophicated situations; 5, occurring in highly eutrophicated situations) according to Nimis (2016)

From: Lichens and other lithobionts on the carbonate rock surfaces of the heritage site of the tomb of Lazarus (Palestinian territories): diversity, biodeterioration, and control issues in a semi-arid environment

Lichen species

Growth form

Eut

Av. frequency per arch. el. (% of quadrats)

Av. frequency per plot (% of quadrats)

Maximum cover (%)

(i)

(ii)

(iii)

(i)

(ii)

(iii)

(i)

(ii)

(iii)

Bagliettoa baldensis (A. Massal.) Vězda

Cr. en.

1

50.0

16.7

13.8

0.8

20

+

Calogaya cfr. decipiens (Arnold) Arup, Frödén, & Søchting§

Cr. ep.

4–5

10.0

16.7

0.4

7.4

+

2

Calogaya pusilla (A. Massal.) Arup, Frödén, & Søchting§

Cr. ep.

2–3

50.0

28.0

1

Candelariella aurella (Hoffm.) Zahlbr.

Cr. ep.

2–4

80.0

33.3

24.6

8.2

18

1

Circinaria calcarea (L.) A. Nordin, Savić, & Tibell

Cr. ep.

2–3

20.0

33.3

25.0

5.0

15.6

2.0

6

16

+

Circinaria contorta (Hoffm.) A. Nordin, Savić, & Tibell

Cr. ep.

3–5

20.0

16.7

9.2

0.8

15

+

Flavoplaca polycarpa (A. Massal.) Arup, Frödén, & Søchting§

Cr. ep.

1–3

20.0

2.9

+

Lepraria sp.

Cr. ep.

60.0

16.7

27.5

1.6

20

1

Myriolecis albescens (Hoffm.) Sliwa, Zhao Xin, & Lumbsch

Cr. ep.

3–4

40.0

16.7

25.0

7.1

6.6

5.0

2

6

+

Myriolecis gr. dispersa (Pers.) Sliwa, Zhao Xin, & Lumbsch

Cr. ep.

2–4

10.0

66.7

25.0

1.3

24.6

2.0

+

3

+

Myriolecis pruinosa (Chaub.) Sliwa, Zhao Xin, & Lumbsch

Cr. ep.

2–4

30.0

50.0

12.1

10.7

2

4

Physconia muscigena (Ach.) Poelt

Fol.

2–4

30.0

16.7

9.2

0.8

+

+

cfr. Pyrenodesmia erodens (Tretiach, Pinna, & Grube) Søchting, Arup, & Frödén§

Cr. en.

1–2

16.7

25.0

9.8

4.0

2

+

Squamarina cfr. gypsacea (Sm.) Poelt

Sq.

1–3

16.7

1.6

+

Variospora aurantia (Pers.) Arup, Frödén, & Søchting§

Cr. ep.

3–4

30.0

100.0

25.0

12.1

63.9

4.0

12

23

+

Verrucaria ochrostoma (Leight.) Trevis.

Cr. en.

4–5

70.0

83.3

43.3

41.0

20

12

Xanthocarpia lactea (A. Massal.) A. Massal.§

Cr. en.

2–3

70.0

66.7

12.1

32.8

1

4

  1. Average frequency per arch. el. ≥ 50%, average frequency per plot ≥ 10%, maximum cover ≥ 20% are reported in italic. Names of species displaying these abundance levels on at least one arch. el. are reported in bold
  2. §Species included in the genus Caloplaca before the taxonomic revision by Arup et al. (2013)